清朝乾隆年间，美丽端庄的姑娘夏紫薇和丫环金锁从济南大明湖来至京城，因在大街拦截宗人府梁大人的轿乘而被殴打，正在市集卖艺的小燕子、柳家兄妹拔刀相助，由此和紫薇成为好友。在此之后，紫薇主仆与小燕子结伴卖艺，经历了无数次刺激的冒险，还结实洋人班杰明。随着友情的加深，紫薇向小燕子吐露了身世，原来她竟是当今圣上的亲生女儿。为了帮紫薇与父皇相认，小燕子鼎力相助，结果却阴差阳进宫错成了还珠格格。调皮的小燕子、端...
永琪认为任务已完成一半，孩子将是他留给额娘的希望。班杰明醒来，将趴在床边睡着的小燕子画下。欣荣生下儿子，永琪即向皇上辞行。皇上尊重其选择，并称若他三年未归，将宣告其去世。永琪带着新写的信和燕子钟，在孩子满月之时孤身骑马出宫，尔康和已有身孕的紫薇在半路等候相送。小燕子和永琪在早已超过约定期限的某日终于…
紫薇向令妃磕头，感谢她多年的照顾。才子美女再次“气儿力顶”，送别紫薇。洞房中，紫薇与尔康以身上的疤痕为引，展开一番追忆与告白。一只山羊离群，小燕子追至山林而迷路。突降大雨，小燕子躲至山洞，又冷又饿又怕。班杰明冒雨赶来，二人顿时相拥。班杰明脱下衣服给小燕子穿，自己着了凉，回家后咳嗽一夜并发烧，小燕子尽…
晴儿和箫剑在洱海以白族风俗举行了婚礼，小燕子班杰明现场献唱。桂嬷嬷给欣荣出主意，让永琪睡前喝下药酒。永琪在意识迷乱中，将欣荣当作小燕子，欣荣接受。小燕子开始练字，给永琪写信。皇上封紫薇为“明珠格格”，并指婚予尔康；太后送来新衣，并送回了福家玉佩。小燕子在放羊时想念永琪，班杰明教她拉小提琴曲“四季”。…
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